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Synonyms: Henslowiaceae Lindl.
Common name: Crypteronia Family.
Number of genera 3. Number of species 11.
Angiosperm. Magnoliopsida.
Disseminule a dehisced fruit, or a seed.
Fruits
Pistil(s) compound; 1; 1-pistillate; with carpels united. Fruit pericarpium, or anthocarp; simple; capsule; loculicidal capsule (of authors, but not Spjut), or ceratium capsule (Crypteronia griffithii C. B. Clarke); capsule not inflated; capsule without operculum; simple; without persistent central column; with styles(s); at apex; not within accessory organ(s), or within accessory organ(s); within receptacle; connate; persistent; more than 1 but less than 10-seeded to many-seeded; 2-seeded (-many); 2–6-carpellate; with carpels united; with carpels remaining united at maturity; with carpels not radiating at maturity; with carpels remaining connected at style; without sterile carpels; apex not beaked; wall woody, or chartaceous; dehiscent. Dehiscent unit seed(s). Dehiscent regularly; passively; and shedding seeds; with replum; fruit without centered partition attached to replum. Epicarp durable; glabrous (without hairs), or not glabrous (with hairs); hairs short; hairs not glandular; without armature; without wing(s); without apical respiratory hole. Mesocarp absent. Endocarp present; not separating from exocarp; thin; not splitting into 1-seeded pyrenes; smooth; without wing; without operculum; without secretory cavities; without mechanism for seedling escape; without grooves; without longitudinal ridges. Funiculus short; short without seed bearing hooks (retinacula); not persisting in fruit after seed shed.
Seeds
Aril absent. Seed minute; less than 1 mm long; 0.5 mm long; straight, or elliptic, or ovate; in transection flattened; not bowl shaped; not nutlike; without winglike beak; without caudate appendage(s); at maturity without food reserves; without canavanine. Sarcotesta absent. Testa present; without fleshy or leathery layer over hard layer; tight; without crease or line separating cotyledons from hypocotyl-radicle; without notch along margin where cotyledons from hypocotyl-radicle tip approach each other; without glands; without bristles; glabrous; without wings, or with wing(s); 2-winged; with wings at both ends; with wings equally developed; with wing(s) fimbriate; with solid wing(s) similar to testa (membranous); without collar; without operculum; colored; monochrome; not becoming mucilaginous when wetted; surrounding embryo. Raphe conspicuous; texture as testa (assumed).
Embryo differentiated from food reserve; well developed; 1 per seed; completely filling testa (no food reserve); chamber basal to wing, or apical to wing, or central to wings, or lateral to wing; at one end of seed not extending into a depression or cup; linear; straight; parallel to seed length; with cotyledons gradually connected to hypocotyl-radicle; without coleorhiza; without simmondsin; without stomata; not green; with 2 or more cotyledons. Cotyledons 2; well developed; 0.5–0.6 times length of embryo; as wide as hypocotyl-radicle; not concealing hypocotyl-radicle; not foliaceous; flat; smooth; with apices entire; with margins separate; basally entire; equal in size; not punctate dotted. Hypocotyl-radicle well developed; straight; not thickened.
Distribution
Old World. Southeastern Asia.
Notes
Cronquist & Mabberley: loculicidal capsule & valves attached & held together by persistent style. Van Beusekon-Osinga et al. (1975) loculicidal capsule. Goldberg: apparently incorrectly - fruit loculicidal, more rarely septicidal or circumscissle capsule, berry. drupe, nut, occ. 3–4-winged. Fruit subglobse to ellipsoid and some of them are puberulous.
Weed information
No USA noxious weeds.
Listed seeds
No ASOA or ISTA listed seeds.
Accepted genera
Axinandra Thwaites -- Crypteronia Blume -- Dactylocladus Oliv.
References specific to this family
Cronquist page 633. Beusekom-Osinga, R.J. van & C.F. van Beusekom. 1975. Delimitation and subdivision of the Crypteroniaceae (Myrtales). Blumea 22:255–266.
General references
Cronquist, A. 1981. An integrated system of classification of flowering plants, 1,262 p. Columbia University Press, New York, Goldberg, A. 1986 (dicots) and 1989 (monocots). Classification, evolution, and phylogeny of the familes of Dicotyledons. Smithsonian Contr. Bot. 58 for dicots (314 pp.) and 71 for monocots (74 pp.). [Goldberg's illustrations are reproduced from older publications and these should be consulted], Gunn, C.R., J.H. Wiersema, C.A. Ritchie, and J.H. Kirkbride, Jr. 1992 and amendments. Families and genera of Spermatophytes recognized by the Agricultural Research Service. Techn. Bull. U.S.D.A. 1796:1–500, Mabberley, D.J. 1987. The plant-book, 706 p. Cambridge University Press, Cambridge, Spjut, R.W. 1994. A systematic treatment of fruit types. Mem. New York Bot. Gard. 70:1–182.
Illustrations
No fruit and poor seed illustrations. Cronquist has no illustration. Seed illustration(s): Karen. Embryo illustration(s): Karen. Karen's plate number and taxon (taxa): 191: Crypteronia griffithii C.B. Clarke (A-B).
• Fruit. 1 of 3. Crypteronia griffithii C. B. Clarke: dehisced fruit with calyx. • Seed. 2 of 3. Crypteronia griffithii C. B. Clarke: seed. • Embryo. 3 of 3. Crypteronia griffithii C. B. Clarke: embryo.
We advise against extracting comparative information from the descriptions. This is much more easily achieved using the DELTA data files or the interactive key, which allows access to the character list, illustrations, full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting or lacking specified attributes, and distributions of character states within any set of taxa. See also Guidelines for using data taken from Web publications.
Cite this publication as: ‘J.H. Kirkbride, Jr., C.R. Gunn, and M.J. Dallwitz. 2000 onwards. Family guide for fruits and seeds: descriptions, illustrations, identification, and information retrieval. Version: 12th April 2021. delta-intkey.com’.
