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Anastrepha and Toxotrypana:
descriptions, illustrations, and interactive keys

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Allen L. Norrbom, Cheslavo A. Korytkowski, Roberto A. Zucchi, Keiko Uramoto, George L. Venable, Jerrett McCormick and Michael J. Dallwitz

Anastrepha dryas Stone

Body. Setae orange brown, or red brown, or dark red brown.

Head. Face with ventral part gradually tapered laterally.

Thorax. Mesonotum length 2.7–3.3 mm (3.0, n=1, Stone 1942b). Postpronotal, presutural supra-alar, dorsocentral, intra-alar and scutellar setae well developed, subequal to or longer than scutellum length. Scutum with 3 (both medial and sublateral) pale postsutural vittae. Scutum posteriorly without brown or orange brown markings (??confirm, check for state 2). Scutum without brown vittae. Scutellum entirely yellow or with dark markings only on extreme base of disk. Mesopleuron mostly yellow to orange, without brown markings. Subscutellum entirely yellow to orange. Mediotergite entirely yellow to orange.

Wings. Wing length 7–8 mm (7.5, n=1, Stone 1942b). Wing pattern typical Anastrepha pattern (S-band complete or at most interrupted at crossvein r-m, C-band and at least proximal arm of V-band present). C-band broadly extending to vein M in cell br along cell bm; covering base of cell r2+3; yellow or orange area posterior to pterostigma broad, extending distally into cells r1 and r2+3 at least to level of midlength of pterostigma. C-band and S-band separated (probable variation), or connected (narrowly along vein R4+5 and sometimes along vein R2+3, cell r1 with basomarginal hyaline spot). Basal hyaline area between C-band and S-band extended to vein R4+5. Cell r1 basomarginal hyaline spot triangular to quadrate. Cell r1 basomarginal hyaline spot apex aligned proximal to crossvein r-m. S-band extended anteriorly to vein R4+5 and covering all of crossvein r-m. Cell bm entirely hyaline or infuscated only along subapical fold. S-band posterior margin without incision in cell cu1. S-band base without extension in middle of cell cu1 to posterior wing margin; without extension in cell a1 to or almost to posterior margin. S-band middle section predominantly or entirely orange, often with brown margins. Subapical hyaline area in radial cells distal to r-m extending anteriorly to vein R2+3. S-band distal section without marginal hyaline band or spots in cell r2+3 or near apices of R2+3 or R4+5. S-band distally not extended to apex of vein M. V-band proximal arm as dark as apical half of S-band; extending more than 1/3 distance from apex of vein Cu1 to apex of vein A1+Cu2; not connected anteriorly to S-band, or connected anteriorly to S-band along vein R4+5 or in cell r2+3 (narrowly, if connected); not connected to S-band in cell dm. V-band distal arm complete; connected to proximal arm of V band. Area surrounding apex of lobe of cell bcu with microtrichia similar in density to area anterdistal to it along vein Cu1. Vein R2+3 without accessory vein. Crossvein dm-cu orientation with anterior end more distal than posterior end.

Abdomen. Abdomen ovate or parallel-sided, syntergite 1+2 gradually broadening or parallel-sided. Abdominal tergite without brown markings. Oviscape length 3.5–4.2 mm (3.86, n=1, Stone 1942b); ratio (oviscape length/mesonotum length) 1.1–1.5 (1.29, n=1, Stone 1942b). Eversible membrane with dorsobasal denticles all sclerotized, in continuous triangular to semicircular or suboval pattern. Eversible membrane with 75–95 denticles (long slender hooklike denticles in 6–8 irregular rows). Aculeus length 3.2–3.9 mm (3.53, n=1, Stone 1942b). Aculeus length/oviscape length 0.91, n=1, Stone 1942b. Aculeus tip length 0.35–0.45 mm (0.40, n=1, Stone 1942b); lateral margins not curved dorsally; not flared outward at or proximal to base; without ridges or lobes; without elongate dorsolateral depressions apically; with fine serrations, or with medium sized serrations; serrated part 0.7–0.85 times length of tip (0.80, n=1, Stone 1942b). Aculeus tip serrations not extending onto dorsal side basally. Aculeus tip with serrations separated by less than width of serration.

Miscellaneous. Data source: Stone 1942b. Data recording inadequate. Sex of recorded specimens: male and female. Species group: pseudoparallela group.

Biology and economic significance. This species may have some economic significance as a pest of fruits of species of Passiflora (Passifloraceae). The main damage is caused by the larvae, which feed inside the fruit. Refer to the 'Tephritidae Databases' in the USDA Compendium of Fruit Fly Host Information (https://coffhi.cphst.org/) for host plant information.

Illustrations. • Habitus, female (dorsal). • Wing. • Terminalia, female.


We advise against extracting comparative information from the descriptions. This is much more easily achieved using the DELTA data files or the interactive key, which allows access to the character list, illustrations, full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting or lacking specified attributes, and distributions of character states within any set of taxa. See also Guidelines for using data taken from Web publications.


Cite this publication as: ‘Norrbom, A.L., Korytkowski, C.A., Zucchi, R.A., Uramoto, K., Venable, G.L., McCormick, J., and Dallwitz, M.J. 2012 onwards. Anastrepha and Toxotrypana: descriptions, illustrations, and interactive keys. Version: 21st November 2023. delta-intkey.com’.

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